This story revolves around the contrast between the "Garnet Prince," a feminized boy who transforms when he falls into darkness while retaining a male heart, and the "Cobalt Princess," a feminized boy with a female heart who transforms when his maiden spirit awakens.

Sharp harbors a desire to live a yuri-filled life as a girl, so he infiltrates an evil organization’s lab, steals a magical potion, and transforms into a woman while keeping his male heart intact. On the other hand, Largo, who possesses the heart of a woman, switches bodies with his childhood friend Forte and begins to awaken to his maiden spirit.